। ॥ ४२१ ॥
In plain words
The body; the whole troop of organs; the breath; the "I"; and the deities presiding over each organ — these five are named as the causes by which every action gets done.
Close to the Sanskrit
The body and the host of organs, the vital breath, the ego, and the presiding deities — these five are declared to be the causes in the accomplishment of all actions.